Help!

Tibor Kercz’s owl trying to climb back on to a branch in Ópusztaszer, Hungary, is overall winner, as well as portfolio winner.
Photograph: Tibor Kercz / Barcroft Images

The laughing dormouse

Andrea Zampatti’s dormouse, taken in Italy, wins the On the Land category.
Photograph: Andrea Zampatti / Barcroft Images

Flying duck

John Threlfal wins the Up in the Air category with his shot of a duck flying in front of a plane trail in Preston, UK.
Photograph: John Threlfal / Barcroft Images

Slap in the face

Troy Mayne wins the Under the Sea category for his sea turtle slapping a passing fish in Bacong, Philippines.
Photograph: Troy Mayne / Barcroft Images

Take me to church

Carl Henry wins highly commended for his trio of penguins heading to church in South Georgia Island, southern Atlantic.
Photograph: Carl Henry / Barcroft Images

Friends in high places

Jean-Jacques Alcalay wins highly commended for his blue wildebeest appearing to ride a wave of hundreds of other wildebeest in Masai Mara, Kenya.
Photograph: Jean-Jacques Alcalay/ Barcroft Images

Great escape

Katy Laveck-Foster wins highly commended for her monkeys enjoying a moped ride in North Sulawesi, Indonesia
Photograph: Katy Laveck Foster / Barcroft Images

Because I’m happy

Penny Palmer wins highly commended for her happy sea otter in California, US.
Photograph: Penny Palmer / Barcroft Images

Hitch a ride

Daisy Gilardini wins highly commended for her polar bear cub clambering on to its mum in Manitoba, Canada.
Photograph: Daisy Gilardini / Barcroft Images

Mudskippers got talent

Daniel Trim wins highly commended for his two mudskippers fighting over territory in Krabi, Thailand.
Photograph: Daniel Trim / Barcroft Images

Outfoxed

Douglas Croft wins highly commended for his fox appearing to use a golf course as his personal toilet in San Jose, US.
Photograph: Douglas Croft / Barcroft Images

What’s up doc?

Olivier Colle’s wild rabbit in Flanders, Belgium, wins highly commended
Photograph: Olivier Colle / Barcroft Images

Caught in the act

Bence Mate wins highly commended for his shot of two bears having some night-time fun in Harghita county, Romania
Photograph: Bence Mate / Barcroft Images

Say what?!

George Cathcart wins highly commended for his shocked seal in San Simeon, California
Photograph: George Cathcart / Barcroft Images